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Ackerman, J.D., Phillips, R.D., Tremblay, R.L., Karremans, A.P., Reiter, N., Peter, C.I., Bogarín, D., Pérez-Escobar, O.A., Liu, H., 2023. Beyond the various contrivances by which orchids are pollinated: global patterns in orchid pollination biology. Botanical J. Linnean Society 202(3): 295–324.

Chinchilla, I.F., Karremans, A.P. & Blanco, M. 2023. Malaxis dressleriana, a new orchid species honoring a botanist for all seasons, Robert Louis Dressler. Orchids (Bull. Amer. Orch. Soc.) 92(8): 608–613.

Gutiérrez Morales, N., Moreno, J. S., Karremans, A.P., & Gil-Amaya, K. 2023. Restrepia santanderensis (Orchidaceae: Pleurothallidinae), a new species from the western slope of the eastern Andes in Colombia. Phytotaxa 598(4): 293–300.

Jost, L., Yanez-Muñoz, M.H., Brito, J., Reyes-Puig, C., Reyes-Puig, J.P., Guayasamín, J.M., Ron, S.R., Quintana, C., Iturralde, G., Baquero, L., Monteros, M., Freire-Fierro, A., Fernández, D., Mendieta-Leiva, G., Morales, J.F., Karremans, A.P., Vázquez-García, J.A., Salazar, G.A., Hágsater, E., Solano, R., Fernández-Concha, G.C. & Arana, M. 2023. Eponyms are important tools for biologists in the Global South. Nat Ecol Evol 7: 1164–1165.

Karremans, A.P. 2023. Karma, a new genus in the Pleurothallidinae (Orchidaceae). Harvard Papers in Botany 28(1): 61-69.

Karremans, A.P. & Jiménez, E. 2023. Pleurothallis pugio (Orchidaceae), a new Pleurothallidinae from Costa Rica. Phytotaxa 599 (3): 150–154.

Karremans, A.P. & Pupulin, F. 2023. Proposal to conserve the name Vanilla (Orchidaceae) with a conserved type. Taxon 72(5): 1121–1122.

Karremans, A.P., Chinchilla, I.F. & Rojas-Alvarado, G. 2023. Studies on Costa Rican Vanilloideae: The return of Cleistes rosea and a reaffirmation of Epistephium ellipticum. Lankesteriana 23(3): 613–622.
http://dx.doi.org/ 10.15517/lank.v23i3.58136

Karremans, A.P., Moreno, J.S., Gil-Amaya, K., Guitérrez Morales, N., Espinosa, F., Mesa, S., Restrepo, E., Rincón-González, M., Serna, A., Sierra-Ariza, M. & Vieira-Uribe, S. 2023. Colombian Orchidaceae: A catalogue of the Pleurothallidinae. Lankesteriana 23(2): 181–400.

Karremans, A.P., Rojas-Alvarado, G. & Chinchilla, I.F. 2023. Epidendrum scandens: an unusual new orchid species endemic to the Cordillera de Talamanca in Costa Rica. Phytotaxa 626(4): 290–296.

Karremans, A.P., Watteyn, C., Scaccabarozzi, D., Pérez-Escobar, O.A. & Bogarín, D. 2023. Evolution of seed dispersal modes in the Orchidaceae: has the Vanilla mystery been solved? Horticulturae 9: 1270.

Perdomo, O., Trujillo Trujillo, E. & Karremans, A.P. 2023. Masdevallia leonor-baeziana (Pleurothallidinae): a new species from the Andean-Amazonian foothills of Caquetá, Colombia. Lankesteriana 23(2): 139–144.

Pupulin, F., Bogarín, D. & Karremans, A.P. 2023. The Lankester Catalogue of Costa Rican Orchidaceae. Lankesteriana 23(Supplement): 1–254.

Solano, R. & Karremans, A.P. 2023. A review of Acianthera subgenus Brenesia (Orchidaceae, Pleurothallidinae), with a new species from Mexico. Acta Botánica Mexicana 130: e2225.

Watteyn, C., Reubens, B., Azofeifa Bolaños, J.B., Solano Campos, F., Pérez Silva, A., Karremans, A.P. & Muys, B. 2023. Cultivation potential of Vanilla crop wild relatives in two contrasting land use systems. European Journal of Agronomy 149: 126890

Watteyn, C., Scaccabarozzi, D., Muys, B., Reubens, B., Ackerman, J.D., Fernández Otárola, M., Guizar Amador, M.F., Karremans, A.P. 2023. Sweet as Vanilla hartii: Evidence for nectar rewards in Vanilla (Orchidaceae) flowers. Flora 303(2023): 152294.


Chinchilla, I.F., Karremans, A.P., & Blanco, M.A. 2022. A new species and a new record of Malaxis (Malaxidinae) from Costa Rica. Lankesteriana 33(1): 37-51.

Karremans, A.P., Bogarín, D., Fernández Otárola, M., Sharma, J., Watteyn, C., Warner, J., Rodríguez Herrera, B., Chinchilla, I.F., Carman, E., Rojas Valerio, E., Pillco Huarcaya, R. & Whitworth, A. 2022. First evidence for multimodal animal seed dispersal in orchids. Current Biology

Pessoa, E. & Karremans, A.P. 2022. A New Masdevallia (Pleurothallidinae: Orchidaceae) from the White-Sand Forests of the Central Amazon, Brazil. Systematic Botany 47(1): 293-298.

Govaerts, R., Karremans, A.P. & Shaw, J. 2022. What’s the date? The Orchid Review 130 (4): 54–59.


Chambers, A., Cibrián-Jaramillo, A., Karremans, A.P., Moreno Martinez, D., Hernandez-Hernandez, J., Brym, M., Resende, M.F.R, Moloney, R., Sierra, S.N., Hasing, T. Alomia, Y.A. & Hu, Y. 2021. Genotyping-By-Sequencing diversity analysis of international Vanilla collections uncovers hidden diversity and enables plant improvement. Plant Science 311: 111019. https://doi.org/10.1016/j.plantsci.2021.111019

Karremans, A.P. 2021. With great biodiversity comes great responsibility: the underestimated diversity of Epidendrum (Orchidaceae). Harvard Papers in Botany 26(2): 299-369. https://doi.org/10.3100/hpib.v26iss2.2021.n1

Karremans, A.P., Rojas-Alvarado, G., Bezerra-Silva, L.E., Flores-Rojas, J., Murillo-Murillo, J.M., Romero-Ceciliano, M. & Segura-Bermúdez, O.A. 2021. Nomenclatural notes in the Pleurothallidinae (Orchidaceae): miscellaneous part II. Phytotaxa 496(2): 134-146. https://doi.org/10.11646/phytotaxa.496.2.3

Ponert, J. Portilla Andrade, M., Karremans, A.P., Chumová, Z. & Trávnícek. 2021. A new species of Pabstiella (Pleurothallidinae, Orchidaceae) from Ecuador. Phytotaxa 500(2): 108-116. https://doi.org/10.11646/phytotaxa.500.2.3

Rojas-Alvarado, G., Blanco, M.A. & Karremans, A.P. 2021. A taxonomic synopsis and morphological characterization of Myoxanthus (Orchidaceae: Pleurothallidinae). Phytotaxa 507(3): 211-258. https://doi.org/10.11646/phytotaxa.507.3.1

Watteyn, C., Scaccabarozzi, D., Muys, B., van der Schueren, N., van Meerbeek, K., Guizar Amador, M.F., Ackerman, J.D., Cedeño Fonseca, M., Chinchilla, I.F, Reubens, B., Pillco Huarcaya, R. Cozzolino, S. & Karremans, A.P. 2021. Trick or Treat? Pollinator attraction in Vanilla pompona (Orchidaceae). Biotropica

Watteyn, C., Dejonghe, O., van Hoyweghen, K., Azofeifa Bolaños, J.B., Karremans, A.P., Vranken, L., Reubens, B., Muys, B. & Maertens, M. 2021. Exploring farmer preferences towards innovations in the vanilla supply chain. Journal of Cleaner Production 330: 129831. https://doi.org/10.1016/j.jclepro.2021.129831


Karremans, A.P., Chinchilla, I.F., Rojas-Alvarado, G., Cedeño-Fonseca, M. Damián, A. & Léotard, G. 2020. A reappraisal of Neotropical Vanilla. With a note on taxonomic inflation and the importance of alpha taxonomy in biological studies. Lankesteriana 20(3): 395-497. https://doi.org/10.15517/lank.v20i3.45203

Karremans, A.P., Salguero, G. Bogarín, D. Oses, L. & Cedeño-Fonseca, M. 2020. Illustrations and studies in Neotropical Orchidaceae: the Specklinia brighamii group (Pleurothallidinae) in Costa Rica. Phytotaxa 447(1): 16–30.

Pupulin, F. & Karremans, A.P. 2020. A new and unusual species of Dichaea (Orchidaceae: Zygopetalinae) from Costa Rica. Blumea 65: 61-64.

Pupulin, F., Belfort-Oconitrillo, N., Karremans, A.P. & Bogarín, D. 2020. Florae costaricensis subtribui Pleurothallidinis Prodromus – Systematics of Echinosepala (Orchidaceae). Harvard Papers in Botany 25(2): 155-190.

Pupulin, F., Bogarín, D. & Karremans, A.P. 2020. Una nueva especie de Cischweinfia (Orchidaceae: Oncidiinae) de Costa Rica, con hábito y flores muy pequeñas. Orquideología 37(1): 4-26.

Reina-Rodríguez, G.A., Karremans, A.P., López-Machado, F. & Cruz-Salcedo, J. 2020. A new Stelis (Orchidaceae: Pleurothallidinae) from the Western Andes of Colombia. Lankesteriana 20(2): 151–158.

Rincón-González, M., Sierra-Ariza, M.A. & Karremans, A.P. 2020. A new species of Acianthera (Orchidaceae, Pleurothallidinae) from the Cordillera Central of Colombia. Phytotaxa 474(1): 093-098. https://doi.org/10.11646/phytotaxa.474.1.10

Rojas-Alvarado, G. & Karremans, A.P. 2020. Revision of the Costa Rican species of Myoxanthus (Pleurothallidinae: Orchidaceae). Phytotaxa 448(1): 001-070. https://doi.org/10.11646/phytotaxa.448.1.1

Salguero Hernández, G., Karremans, A.P., Pupulin, F. & Strigari, S. 2020. The New Refugium Botanicum. Arpophyllum giganteum. Orchids (Bull. Amer. Orch. Soc.) 89(10): 766–768.

Watteyn, C., Fremout, T., Karremans, A.P., Pillco Huarcaya, R., Azofeifa Bolaños, J.B., Reubens, B.& Muys, B. 2020. Vanilla distribution modeling for conservation and sustainable cultivation in a joint land sparing/sharing concept. Ecosphere 11(3):e03056.


Bogarín, D., Fernández, F., Karremans, A.P., Pupulin, F., Smets, E. & Gravendeel, B. 2019. Floral anatomy and evolution of pollination syndromes in Lepanthes and close relatives. In: A. M. Pridgeon & A. R. Arosemena (Eds.), Proceedings of the 22nd World Orchid Conference, vol. 1 (pp. 396‒410). Guayaquil, Ecuador: Asociación Ecuatoriana de Orquideología.

Bogarín, D., Pérez-Escobar, Karremans, A.P., Fernández, M., Kruizinga, J., Pupulin, F., Smets, E. & Gravendeel, B. 2019. Phylogenetic comparative methods improve the selection of characters for generic delimitations in a hyperdiverse Neotropical orchid clade. Scientific Reports 9:15098. https://doi.org/10.1038/s41598-019-51360-0

Karremans, A.P. 2019. To be or not to be a Stelis. Lankesteriana 19(3): 281-343. https://doi.org/10.15517/lank.v19i3.40082

Karremans, A.P. & Díaz-Morales, M. 2019. The Pleurothallidinae: extremely high speciation driven by pollinator adaptation. In: A. M. Pridgeon & A. R. Arosemena (Eds.), Proceedings of the 22nd World Orchid Conference, vol. 1 (pp. 363‒388). Guayaquil, Ecuador: Asociación Ecuatoriana de Orquideología.

Karremans, A.P., Aguilar-Sandí, D., Artavia-Solís, M., Cedeño-Fonseca, M., Chinchilla, I. F., Gil, K., Rojas-Alvarado, G., Solano-Guindon, N. & Villegas- Murillo, J. 2019. Nomenclatural notes in the Pleurothallidinae (Orchidaceae): miscellaneous. Phytotaxa 406(5), 259‒270. https://doi.org/10.11646/phytotaxa.406.5.1

Thoerle, L. & Karremans, A.P. 2019. The genus Muscarella (Orchidaceae): two new synonyms. Phytotaxa 395(1): 48-50. https://doi.org/10.11646/phytotaxa.395.1.6


Bogarín, D, A. Karremans, A.P. & Fernández, M. 2018. Genus-level taxonomical changes in the Lepanthes affinity (Orchidaceae, Pleurothallidinae). Phytotaxa 340 (2): 128-136. https://doi.org/10.11646/phytotaxa.340.2.2

Bogarín, D, Pérez-Escobar, OA, Groenenberg, D, Holland, SD, Karremans, A.P., Lemmon, EM, Lemmon, AR, Pupulin, F, Smets, E, Gravendeel, B. 2018. Anchored hybrid enrichment generated nuclear, plastid and mitochondrial T markers resolve the Lepanthes horrida (Orchidaceae: Pleurothallidinae) species complex. Molecular Phylogenetics and Evolution 129(2018): 27-47. https://doi.org/10.1016/j.ympev.2018.07.014

Cedeño-Fonseca, M., Karremans, A.P. & O. O. Ortiz. 2018. Monstera limitaris (Araceae) a new species from the border between Costa Rica and Panama. Phytotaxa 376(1): 037–042. https://doi.org/10.11646/phytotaxa.376.1.4

Karremans, A.P. & Jiménez, JE. 2018. Pleurothallis hawkingii and Pleurothallis vide-vallis (Orchidaceae; Epidendroideae), two new species from Cordillera de Guanacaste in Costa Rica. Phytotaxa 349(2): 185-191. https://doi.org/10.11646/phytotaxa.349.2.10

Karremans, A.P. & Lehmann, CP. 2018. A highly threatened new species of Vanilla from Costa Rica. Lindleyana in Orchids (Bull. Amer. Orch. Soc.) 87(4): 304–307.

Reina-Rodríguez, G.A. & Karremans, A.P. 2018. A new Platystele (Orchidaceae, Pleurothallidinae) from the Western mountains of Colombia. Phytotaxa 375(3): 235–242. https://doi.org/10.11646/phytotaxa.375.3.6

Vieira-Uribe, S. & Karremans, A.P. 2018. La Specklinia (Orchidaceae) con señuelo, una asombrosa nueva especie de los Andes occidentales en Colombia. Orquideología 32(2): 164–168.


Collantes, B. & Karremans, A.P. 2017. An eccentric new species of Stelis from Peru (Orchidaceae: Pleurothallidinae). Brittonia 69(2): 218–221. https://doi.org/10.1007/s12228-017-9462-8

Karremans, A.P. & Bogarín, D. 2017. Two novelties in Platystele (Ochidaceae: Pleurothallidinae) from Costa Rica. Lankesteriana 17(2): 215-221. https://doi.org/10.15517/lank.v17i2.30044

Karremans, A.P. & Davín, N. 2017. Genera Pleurothallidinarum: The Era of Carl Luer. Lankesteriana 17(2): I-VIII. https://doi.org/10.15517/lank.v17i2.30273

Karremans, A.P. & Díaz-Morales, M. 2017. Novelties in Costa Rican Stelis (Ochidaceae: Pleurothallidinae): Two new species and a new record in the “Dracontia group”. Lankesteriana 17(2): 193–202. https://doi.org/10.15517/lank.v17i2.29928

Karremans, A.P., Rodríguez-Martínez, L & Rincón-Useche, C. 2017. Four new species of Pleurothallis (Orchidaceae: Pleurothallidinae) from the Valle del Cauca department in Colombia. Lankesteriana 17(2): 251-260. https://doi.org/10.15517/lank.v17i2.30155

Moreno, JS, Vieira-Uribe, S & Karremans, A.P. 2017. A new species of Lepanthes (Orchidaceae: Pleurothallidinae) from Colombia with a large protruding column. Lankesteriana 17(2): 227-234. https://doi.org/10.15517/lank.v17i2.30076

Pérez-Escobar, OA, Chomicki, G, Condamine, FL, Karremans, A.P., Bogarín, D, Matzke, NJ, Silvestro, D & Antonelli, A. 2017. Recent origin and rapid speciation of Neotropical orchids in the world’s richest plant biodiversity hotspot. New Phytologist 215: 891–905. https://doi.org/10.1111/nph.14629

Pupulin, F & Karremans, A.P. 2017. Horichia dressleri. A new genus recorded for Costa Rica and a truly rare orchid? Lindleyana in Orchids (Bull. Amer. Orch. Soc.) 86(8): 610–617.

Pupulin, F, Karremans, A.P. & Belfort Oconitrillo, N, 2017. Two new species of Echinosepala (Orchidaceae: Pleurothallidinae). Lankesteriana 17(2): 285–304. https://doi.org/10.15517/lank.v17i2.30205

Pupulin, F, Karremans, A.P. & Strigari, S. 2017. The New Refugium Botanicum. Ida (Sudamerlycasteandreettae. Orchids (Bull. Amer. Orch. Soc.) 86(7): 496–498.

Rojas-Alvarado, G & Karremans, A.P. 2017. Additions to the Costa Rican Myoxanthus (Orchidaceae: Pleurothallidinae). Lankesteriana 17(2): 203-214. https://doi.org/10.15517/lank.v17i2.29933

Vieira-Uribe, S. & Karremans, A.P. 2017. A new species of Andinia subgen. Brachycladium (Orchidaceae: Pleurothallidinae) from Colombia. Lankesteriana 17(2): 311-314. https://doi.org/10.15517/lank.v17i2.30210

Wilson, M, Frank, GS, Jost, L, Pridgeon, AM, Vieira-Uribe, S & Karremans, A.P. 2017. Phylogenetic analysis of Andinia (Pleurothallidinae; Orchidaceae) and a systematic re-circumscription of the genus. Phytotaxa 295(2): 101–131. https://doi.org/10.11646/phytotaxa.295.2.1


Bogarín, D & Karremans, A.P. 2016. A new Brachionidium (Orchidaceae: Pleurothallidinae) from the first botanical expedition to the Volcán Cacho Negro, Costa Rica. Systematic Botany 41(4): 919-923. https://doi.org/10.1600/036364416X694017

Bonilla Morales, MM, Aguirre Morales, AC, Mosquera Hernández, JJ & Karremans, A.P. 2016. Distribution of Platystele sylvestrei (Orchidaceae: Pleurothallidinae): first records for Colombia. Rev. Biodivers. Neotrop.  6(2): 131-134. https://doi.org/10.18636/bioneotropical.v6i2.354

Chiron, GR, Karremans, A.P. & van den Berg, C. 2016. Nomenclatural notes in the Pleurothallidinae (Orchidaceae): Phloeophila. Phytotaxa 270: 56–62. https://doi.org/10.11646/phytotaxa.270.1.6

Damián, A & Karremans, A.P. 2016. A New Species of Stelis (Orchidaceae: Pleurothallidinae) from Peru. Systematic Botany 41(2): 293-297. https://doi.org/10.1600/036364416X691876

Díaz-Morales, M & Karremans, A.P. 2016. Epidendra Nova Talamancana. Phytotaxa 272(4): 248-256. https://doi.org/10.11646/phytotaxa.272.4.2

Díaz-Morales, M & Karremans, A.P. 2016. The New Refugium Botanicum. Specklinia endotrachys. Orchids 85(5): 332-334.

Karremans, A.P. 2016. Genera Pleurothallidinarum: an updated phylogenetic overview of Pleurothallidinae. Lankesteriana 16(2): 219-241. https://doi.org/10.15517/lank.v16i2.26008

Karremans, A.P. 2016. Editorial: Lankesteriana a no impact journal… or is it?. Lankesteriana 16(1): I-III.

Karremans, A.P., Albetazzi, FJ, Bakker, FT, Bogarín, D, Eurlings, MCM, Pridgeon, A, Pupulin, F & Gravendeel, B. 2016. Phylogenetic reassessment of Specklinia and its allied genera in the Pleurothallidinae (Orchidaceae). Phytotaxa 272(1): 1-36. https://doi.org/10.11646/phytotaxa.272.1.1

Karremans, A.P., Bogarín, D, Díaz-Morales, M, Fernández, M, Oses, L & Pupulin, F. 2016. Phylogenetic reassessment of Acianthera (Orchidaceae: Pleurothallidinae). Harvard Pap. Bot. 21(2): 171-187. https://doi.org/10.3100/hpib.v21iss2.2016.n4

Karremans, A.P., van Heuven, B, Langelaan, R & Gravendeel, B. 2016. Documentation of floral secretory glands in Pleurothallidinae (Orchidaceae) using Scanning Electron Microscopy (SEM). Bio-protocol 6(22): e2021. https://doi.org/10.21769/BioProtoc.2021

Karremans, A.P. & Vieira-Uribe, S. 2016. Andinia lueri, una nueva especie confundida con A. dactyla (Orchidaceae: Pleurothallidinae). Orquideología XXXIII: 112-122.

Oses, L & Karremans, A.P. 2016. A note on Masdevallia zahlbruckneri and M. utriculata. Harvard Pap. Bot. 21(2): 325-336. https://doi.org/10.3100/hpib.v21iss2.2016.n13


Bogarín, D & Karremans, A.P. 2015. Prosthechea tinukiana sp. nov. (Orchidaceae: Laeliinae): an update of the Prosthechea prismatocarpa complex. Turk. J. Bot. 39: 499-505. https://doi.org/10.3906/bot-1408-38

Bogarín, D., Karremans, A.P. & Muñoz, M. 2015. Brachionium kirbyi, a new species honoring the founder of the “Bosque de Paz” Orchid Project in Costa Rica. Die Orchidee 66(5): 404-409.

Dressler, R.L. & Karremans, A.P. 2015. Sobralia rogersiana, un monstruo maravilloso. Orquideología 32(1): 30-37.

Karremans, A.P. 2015. Visualizing Pleurothallids: Lankesteriana. Lindleyana in Orchids (Bull. Amer. Orch. Soc.) 84(5): 304-312.

Karremans, A.P., Bogarín, D, Pupulin, F, Luer, CA, & Gravendeel, B. 2015. The glandulous Specklinia: morphological convergence versus phylogenetic divergence. Phytotaxa 218(2): 101-127. https://doi.org/10.11646/phytotaxa.218.2.1

Karremans, A.P. & M. Rincón-González. 2015. Nomenclatural notes in the Pleurothallidinae (Orchidaceae): Apoda-prorepentia. Phytotaxa 238(2): 174-182. https://doi.org/10.11646/phytotaxa.238.2.5

Karremans, A.P. 2015. Nomenclatural notes in the Pleurothallidinae (Orchidaceae): Stelis. Phytotaxa 203(3): 292-296. https://doi.org/10.11646/phytotaxa.406.5.1

Karremans, A.P., Bogarín, D & Gravendeel, B. 2015. Specklinia lugduno-batavae (Pleurothallidinae: Orchidaceae), a new species in the S. digitalis group. Blumea 59: 180-184. https://doi.org/10.3767/000651915X687859

Karremans, A.P., Pupulin, F., Grimaldi, D., Beentjes, K.K., Butot, R., Fazzi, G.E., Kaspers, K., Kruizinga, J., Roessingh, P., Smets, E.F.  & Gravendeel, B. 2015. Pollination of Specklinia by nectar-feeding Drosophila: the first reported case of a deceptive syndrome employing aggregation pheromones in Orchidaceae. Annals of Botany 116(3): 437-455. https://doi.org/10.1093/aob/mcv086

Karremans, A.P., Pupulin, F., and Gravendeel, B. 2015. Specklinia dunstervillei, a new species long confused with Specklinia endotrachys (Orchidaceae: Pleurothallidinae). PLoS One 10(7): e0131971. https://doi.org/10.1371/journal.pone.0131971

Rodríguez-Martínez, L, Rincón-Useche, C & Karremans, A.P. 2015. A New Pleurothallis from Colombia, with a Note on Ancipitia and Colombiana (Orchidaceae: Pleurothallidinae). Systematic Botany 40(1): 75-78. https://doi.org/10.1600/036364415X686350


Fernández, M, Bogarín, D, Karremans, A.P. & Jiménez, D. 2014. New species and records of Orchidaceae from Costa Rica III. Lankesteriana 13(3): 259-282. https://doi.org/10.15517/lank.v13i3.14363

Karremans AP. 2014. Lankesteriana, a new genus in the Pleurothallidinae (Orchidaceae). Lankesteriana 13(3): 319-332. https://doi.org/10.15517/lank.v13i3.14368


Bogarín, D & Karremans, A.P. 2013. Trichocentrum pupulinianum (Orchidaceae). Lindleyana in Orchids (West Palm Beach) 82(2): 106–110.

Bogarín, D, Karremans, A.P., Rincón, R & Gravendeel B. 2013. A new Specklinia (Orchidaceae: Pleurothallidinae) from Costa Rica and Panama. Phytotaxa 115(2): 31-41. https://doi.org/10.11646/phytotaxa.115.2.1

Karremans, A.P. & Bogarín, D. 2013. Three New Species of Dracontia (Pleurothallidinae, Orchidaceae) from Costa Rica. Systematic Botany 38(2): 307-315. https://doi.org/10.1600/036364413X666796

Karremans, A.P., Bakker, FT, Pupulin, F, Solano-Gomez, R & Smulders, MJM. 2013. Phylogenetics of Stelis and closely related genera (Orchidaceae: Pleurothallidinae). Plant Syst. Evol. 29(1): 69-86. https://doi.org/10.1007/s00606-012-0712-7

Pupulin, F., AP Karremans & S. Strigari. 2013. Taxonomie in Aquarell: die Speckliniaendotrachys-Gruppe, Teil 1. Taxonomy in watercolor: the Specklinia endotrachys group, part 1. Die Orchidee (Hamburg) 64(5): 392-399.

Pupulin, F., AP Karremans & S. Strigari. 2013. Taxonomie in Aquarell: die Speckliniaendotrachys-Gruppe, Teil 2. Taxonomy in watercolor: the Specklinia endotrachys group, part 2. Die Orchidee (Hamburg) 64(6): 475-485.


Bogarín, D, Pupulin, F & Karremans AP. 2012. Three New Lepanthes (Orchidaceae: Pleurothallidinae) from South-East Costa Rica. Lankesteriana 12(2): 107-114. https://doi.org/10.15517/lank.v0i0.11748

Karremans AP. 2012. Illustrations and Studies in Neotropical Orchidaceae. 3. On the identity of Dracontia pachyglossa and D. ramonensis; with a new species, D. lueriana (Pleurothallidinae). Ann. Naturhist. Mus. Wien, B. 113: 119-132.

Karremans AP, D Bogarín, M Fernández, CM Smith & Blanco, MA. 2012. New species and records of Orchidaceae from Costa Rica. II. Lankesteriana 12(1): 19-51. https://doi.org/10.15517/lank.v12i1.18271

Karremans AP & Smith, CM. 2012. A note on genus Dracontia (Orchidaceae: Pleurothallidinae), with a new species. Harvard Pap. Bot. 17(1): 13-17. https://doi.org/10.2307/41761738

Pupulin, F, Karremans, A.P. & Gravendeel, B. 2012. A reconsideration of the Empusellous species of Specklinia (Orchidaceae: Pleurothallidinae) in Costa Rica. Phytotaxa 63: 1-20. https://doi.org/10.11646/phytotaxa.63.1.1


Fernández, M, Karremans, A.P. & Pupulin, F. 2011. Orchid Pollinaria up close and personal. Orchids 80: 362-363,

Karremans, A.P. 2011. Dracontia, the little-known dragon orchids. Orchids 80(9): 560-566.

Karremans, A.P. & Bogarín, D. 2011. Pleurothallis adventurae (Orchidaceae: Pleurothallidinae), eine neue art aus einer unerforschten region in Costa Rica. Orchideen J. 18(3): 111-114.

Karremans, A.P. & Muñoz, M. 2011. Pleurothallis silvae-pacis (Orchidaceae: Pleurothallidinae): a new species found at the Bosque de Paz Biological Reserve in Costa Rica. Orch. Rev. 121: 155-159.


Bogarín, D & Karremans, A.P. 2010. A new Platystele (Orchidaceae: Pleurothallidinae) from Central Costa Rica / Un nuevo Platystele (Orchidaceae: Pleurothallidinae) de la parte central de Costa Rica. Orquideología 27(2): 208-220.

Karremans, A.P. & Hágsater, E. 2010.Confusion in Epidendrum brenesi Schltr., and a new Costa Rican species: Epidendrum sotoanum (Orchidaceae). Lankesteriana 9(3): 403-409. https://doi.org/10.15517/lank.v0i0.12076

Pupulin, F & Karremans, A.P. 2010. Crossoglossa sotoana (Orchidaceae: Malaxideae), a new species honoring the late Mexican botanist, Miguel Ángel Soto Arenas. Lankesteriana 9(3): 443-446.

Pupulin, F & Karremans, A.P. 2010. Two new species and a new record of Epidendrum (Orchidaceae: Laeliinae) from Costa Rica. Selbyana 30(2): 196.


Karremans, A.P. 2009. Prosthechea madrense, a reconsideration of Epidendrum madrense Schltr.(Orchidaceae: Laeliinae). Acta Botánica Mexicana 88: 47-57.


Bogarín, D, Karremans, A.P. & Pupulin, F. 2008.New species and records of Orchidaceae from Costa Rica. Lankesteriana 8(2): 53-74.

Pupulin, F & Karremans, A.P. 2008.The strange story of the two Hoffmann´s Epidendrums. Lindleyana in Orchids 77(6): 454-457.

Pupulin, F & Karremans, A.P. 2008.The Orchid Pollinaria Collection at Lankester Botanical Garden. Selbyana 29(1): 69-86.

Pupulin, F, Karremans, A.P. & Merino, G. 2008. Trichocentrum loyolicum, a new species in the genus Trichocentrum s.s. Harvard Pap. Bot. 13(2): 219-222.


Pupulin, F & Karremans, A.P. 2007. Prosthechea ionophlebia and P. chacaoensis two different species, but which is which? Lindleyana in Orchids 76(3): 202-208.